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HeLa and HEK 293T cells were maintained in either DMEM medium (Gibco) containing 4.5 g l −1 glucose, 110 mg l −1 pyruvate, 4 mM glutamine, 10% FBS, 1% penicillin/streptomycin or in RPMI medium (Gibco) containing 2 mM glutamine, 10% FBS, 1% penicillin and streptomycin

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The MS was operated in the positive electrospray mode at a capillary temperature of 450 °C, source temperature of 150 °C, capillary voltage of 0.5 kV, desolvation gas flow of 900 L/h, and cone gas flow of 50 L/h